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In short, admixture between two very distinct populations has the concrete result of producing striking genomic correlations.
Over time these correlations dissipate due to recombination.
The authors checked all the various combinations, and these came out at the top as likely admixtures (i.e., the two sources produce particular patterns in the target).
The technical reason that the authors consider is that STRUCTURE based inferences may overestimate admixture when reference populations are not appropriate.
And yet the authors still concede that 800 years is simply difficult to credit when one consults the historical literature.
Please remember that these triads should not be taken literally.
The Uygur are not descended from Japanese and Italians.
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Subsequent mating between peoples of neighboring regions, resulting in isolation-by-distance (LAO et al., 2008; NOVEMBRE et al., 2008).